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He handed over eight muzzled coyotes to an animal shelter, in a scene reminiscent of his glory days as South Korea's top scientist.

Dr Hwang said he had developed a new technique to clone wild animals.

He received a suspended prison term in 2009 for embezzlement after he falsely claimed to have cloned human embryos.

He was once a national hero in South Korea, lavished with government funds and the newly created title of supreme scientist.

He was expected to lead the nation in the lucrative new field of biotechnology.

But key parts of research papers published in leading academic journals in 2004 and 2005 were shown to have been fabricated.

Dr Hwang falsely claimed to have cloned human embryos and created a line of human stem cells that led to hope of cures for a range of degenerative diseases.

Scientists around the world were scandalised by the fraud and the professor was dismissed from his post at Seoul National University and stripped of his titles.

A veterinarian by training, Dr Hwang did have real success cloning animals, including the...